The enormous amount of data now available about the pollen tube clearly reflects its qualities as a biological model that go much beyond that of a carrier of sperm cells essential for plant reproduction the diversity of techniques and methodologies currently used to study pollen and pollen tube. Pollen grains harbour the haploid microgametophytes of seed plants spermatophytes that deliver twin sperm cells to the female gametophyte via the pollen tube pollen contains high levels of stored transcripts and protein for use during germination and pollen tube growth. Here we first survey advances in the cellular and molecular understanding of angiosperm pollen and stigma biology including pollen stigma adhesion pollen hydration and germination and pollen tube emergence and invasion. When the pollen tube reaches an ovule one sperm cell fuses with the haploid egg cell to form the embryo while the second sperm cell fuses with a diploid accessory cell the central cell to form the endosperm a specialized food storage tissue communication between pollen and pistil is an essential factor in many aspects of this process for
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